Two years later, Megatron is the secondary antagonist after his master, the Fallen. Megatron serves as the primary antagonist of the 2007 film. Optimus calls Megatron his brother Cullen interpreted this line literally, but while the writers state this was the case in early drafts, they consider the line in its final context to be metaphorical. The prototype design had a much narrower face than the one seen in the finished film. Fans disliked leaked images of the character's head design, so it was redesigned during filming. Orci and Kurtzman also felt it would be "the equivalent to Darth Vader his own lightsaber and having someone else swing him around". The deviation from Megatron's 1980s Walther P38 pistol form was done to avoid morphing. Weaving recorded his lines in Australia, directed by Bay through iChat. Weaving's voice had been the inspiration Bay gave to his animators for the character, and the character's voice had become that of Weaving in the director's mind. Bay considered his voice too soft and felt it would be disrespectful to ask Welker to change it. Frank Welker, who played the part on television opposite Peter Cullen, auditioned to reprise the role. When he is defrosted, he keeps his original alien jet form out of vanity. Megatron crash-landed in the Arctic years ago in his quest to obtain the All Spark. Hugo Weaving voices Megatron, the main Decepticon antagonist, leader of the Decepticons, a master of warfare and a disciple of the Fallen. At some point, Leonard Nimoy and Frank Welker (the original voice of Megatron) were considered for voicing the Fallen, before Todd came in (who had previously worked with director Michael Bay on The Rock). The writers selected The Fallen after looking through various cartoons and comics for new characters, because he was the most "elemental" villain. Although he does not transform in the film, his toy versions transform into a "Cybertronian destroyer" aircraft. He is capable of opening Space Bridges at will. During this time, he was able to communicate with the Decepticons via an interdimensional window, and it was the image of his face, seen through this wind, that served as inspiration for the Decepticon insignia. Although in the film, he is initially shown to be in the Nemesis in some kind of stasis for an unexplained reason, the prequel novels for the film say the Fallen was imprisoned in another dimension thousands of years ago by his brethren as a punishment for his betrayal.
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Tony Todd voices The Fallen, the true leader of the Decepticons who was one of the Dynasty of Primes and Megatron's master and the main antagonist of the 2009 film Transformers: Revenge of the Fallen.